Preprints of submitted work (not peer-reviewed) can be found here.
Green open access Author Accepted Versions can be found by following the “OA version” link to the University of Hull repository, Hydra. Gold open access publications are indicated by the “open access” link.

Peer-reviewed publications


  1. Ward, AJW, Schaerf, T, Herbert-Read, JE, Morrell, LJ, Sumpter, DJT & Webster, M. (in press) Local interactions and global properties of free-ranging stickleback shoals. Royal Society open science 4: 170043 [open access]
  2. Henri, DC, Morrell LJ & Scott, GW (2017) Student perceptions of their autonomy at University. Higher Education 75(3), 507-516 doi:10.1007/s10734-017-0152-y [open access] [blog summary]
  3. Johannesen, A, Dunn, AM & Morrell, LJ. (2017) Disturbed flow can create a sensory refuge for aggregated prey. PeerJ 5:e3121 [open access] [blog summary]
  4. Rothery, L, Scott GS & Morrell LJ. (2017) Colour preferences of UK garden birds at supplementary seed feeders. PLoS ONE 12(2): e0172422 [open access] [blog summary]
  5. 2016

  6. Kelley, JL, Rodgers, GM & Morrell, LJ. (2016) Conflict between background matching and social signalling in a colourful freshwater fish. Royal Society open science 3: 160040 [open access][blog summary]
  7. Hansen, ML, Morrell, LJ & Ward, AJW (2016) The effect of temporally variable environmental stimuli and group size on emergence behavior. Behavioural Ecology 27: 939-945 [link] [OA version]
  8. Kimbell, HS & Morrell, LJ. (2016) Turbidity weakens selection for assortment by body size in groups. Behavioural Ecology 27: 545-552 [link] [OA version]
  9. 2015

  10. Perez, DM, Heatwole, SJ, Morrell, LJ & Backwell, PRY. (2015) Handedness in fiddler crab fights. Animal Behaviour 110: 99-104 [link] [OA version]
  11. Henri, DC, Morrell, LJ & Scott, GW. (2015) As a clearer question, get a better answer. [version 1; 1 approved, 1 approved with reservations] F1000Research 4: 901 [open access]
  12. Kimbell, HS & Morrell LJ. (2015) ‘Selfish herds’ of guppies follow complex movement rules, but not when information is limited. Proceedings of the Royal Society of London Series B 282: 20151558 [link] [OA version]
  13. Morrell, LJ & Joyce DA. (2015) Interactive lectures: clickers or personal devices? [version 1; referees: 2 approved] F1000Research 4: 64 [open access]
  14. Kimbell, HS & Morrell, LJ (2015) Turbidity influences individual and group level responses to predation in guppies (Poecilia reticulata). Animal Behaviour 103:179-185 [link] [OA version]
  15. Rodgers, GM, Downing, B & Morrell, LJ. (2015). Prey body size mediates the predation risk associated with being ‘odd’. Behavioural Ecology 26: 242-246 [link] [OA version]
  16. Morrell, LJ, Greenwood, L & Ruxton, GD. (2015) Consequences of variation in predator attack for the evolution of the selfish herd. Evolutionary Ecology 29: 107-121 [link ] [OA version] [blog summary]
  17. 2014

  18. Ruxton, GD, Humphries, S, Morrell, LJ & Wilkinson, DM. (2014) Why is eusociality an almost exclusively terrestrial phenomenon? Journal of Animal Ecology 83: 1248-1255 [link] [OA version]
  19. Johannesen, A, Dunn, AM & Morrell, LJ. (2014) Prey aggregation is an effective olfactory predator avoidance strategy. PeerJ 2:e408 [open access]
  20.  Morrell LJ. (2014) Use of feed-­forward mechanisms in a novel research­‐led module.  Bioscience Education 22: 70-81 [open access]
  21. 2013

  22. Rodgers, GM, Gladman, NW, Corless, H & Morrell, LJ. (2013) Costs of colour change in fish: food intake and behavioural decisions. Journal of Experimental Biology 216: 2760-2767 [link] [OA version]
  23. Ward, AJW, Herbert-Read, JE, Jordan, LA, James, R, Krause, J, Ma, Q, Rubenstein, DI, Sumpter, DJT & Morrell, LJ. (2013) Initiators, leaders and recruitment mechanisms in the collective movements of damselfish.  American Naturalist 181: 748-760 [link]
  24. Rodgers, GM, Kimbell, H & Morrell, LJ. (2013) Mixed phenotype grouping: the interaction between oddity and crypsis. Oecologia 172: 59-68 [link] [OA version]
  25. Herbert-Read, JE, Krause, S, Morrell, LJ, Shaerf, TM, Krause J & Ward AJW. (2013) The role of individuality in collective group movement. Proceedings of the Royal Society of London Series B 280: 20122564 [link]
  26. 2012

  27. Johannesen, A, Dunn, A & Morrell, LJ. (2012) Olfactory cue use by three-spined sticklebacks foraging in turbid water: prey detection or prey location? Animal Behaviour 84: 151-158 [link] [OA version]
  28. Morrell, LJ, Hentley, WT, Wickens, VJ, Wickens, JB & Rodgers, GM. (2012) Artificial enhancement of an extended phenotype signal increases investment in courtship in three-spine sticklebacks. Animal Behaviour 84: 93-101 [link] [OA version]
  29. Burns, AL, Herbert-Read, JE, Morrell, LJ & Ward, AJW. (2012) Consistency of leading and following in moving shoals of mosquitofish (Gambusia holbrooki). PLoS ONE 7(5): e36567 [open access]
  30. Foster, EA, Franks, DW, Morrell, LJ, Balcomb, KC, Parsons, KM, van Ginnekin, A & Croft, DP. (2012) Social network correlates of food availability in an endangered population of killer whales (Orcinus orca). Animal Behaviour 83: 731-736 [link]
  31. 2011

  32. Kelley, JL, Morrell, LJ, Inskip, C, Krause, J & Croft, DP. (2011) Predation risk shapes social networks in fission-fusion populations. PLoS ONE 6(8): e24280 [open access]
  33. Rodgers, GM, Askwith, B, Ward, J & Morrell, LJ. (2011) Balancing the dilution and oddity effects: decisions depend on body size. PLoS ONE 6(7): e14819 [open access]
  34. Hirsch, B & Morrell, LJ. (2011) Measuring marginal predation in animal groups. Behavioural Ecology 22: 648-656 [link]
  35. Morrell, LJ, Ruxton, GD & James, R. (2011) Spatial positioning in the selfish herd. Behavioral Ecology 22: 16-22 [link] [OA version]
  36. Morrell, LJ, Ruxton, GD & James, R. (2011) The temporal selfish herd. Proceedings of the Royal Society of London Series B 278: 605-612 [link] [OA version]
  37. 2010

  38. Rodgers, GM, Kelley, JL & Morrell, LJ. (2010) Colour change and assortment in the western rainbowfish. Animal Behaviour 79: 1025-1030 [link] [OA version]
  39. Chapman, BB, Morrell, LJ & Krause, J. (2010) Unpredictability in food supply during early life drives the development of boldness in fish. Behavioural Ecology 21: 501-506 [link]
  40. Chapman, BB, Morrell, LJ, Tosh, C, & Krause, J. (2010) Behavioural consequences of sensory plasticity in guppies. Proceedings of the Royal Society of London Series B 277: 1395-1401 [link]
  41. 2009

  42. Chapman, BB, Morrell, LJ & Krause, J. (2009) Plasticity in male courtship behaviour as a function of light intensity in guppies. Behavioral Ecology and Sociobiology 63: 1757–1763 [link]
  43. Ioannou, CC, Morrell, LJ, Ruxton, GD & Krause, J (2009) The effect of prey density on predators: conspicuousness and attack success are sensitive to spatial scale. American Naturalist 173: 499-506 [link]
  44. Dyer, JRG, Croft, DP, Morrell, LJ & Krause, J (2009) Shoal composition determines foraging success in guppies. Behavioral Ecology 20: 165-171 [link]
  45. 2008

  46. Morrell, LJ & Romey, WL (2008) Optimal individual positions in animal groups. Behavioral Ecology 19: 909-919 [link] [OA version]
  47. Botham, MS, Hayward, RK, Morrell, LJ, Croft, DP, Ward JR, Ramnarine, I & Krause, J (2008) Risk sensitive anti-predator behaviour in guppies. Ecology 89: 3174-3185 [link]
  48. Morrell, LJ, Croft, DP, Dyer, JRG, Chapman, BB, Kelley, JL, Laland, KN & Krause, J (2008) Association patterns and foraging behaviour in natural and artificial guppy shoals. Animal Behaviour 76: 855-864 [link] [OA version]
  49. Morrell, LJ & James, R (2008) Mechanisms for aggregation in animals: Rule success depends on ecological variables. Behavioral Ecology 19: 193-201 [link]
  50. Chapman, BB, Morrell, LJ, Benton, TG & Krause, J (2008) Early interactions with adults mediate the development of predator defenses in juvenile guppies Poecilia reticulata. Behavioral Ecology 19: 87-93 [link]
  51. Dyer, JRG, Ioannou, C, Morrell, LJ, Croft, DP, Couzin, ID, Waters, DA, Wong, Y & Krause, J. (2008) Consensus decision making in human crowds. Animal Behaviour 75: 461-470 [link]
  52. Thomas, POR, Croft, DP, Morrell, LJ, Davis, A, Faria, JJ, Dyer, JRG, Piyapong, C, Ramnarine, I, Ruxton, GD & Krause, J. (2008) Does defection during predator inspection affect social structure in wild shoals of guppies? Animal Behaviour 75: 43-55 [link]
  53. 2007

  54. Piyapong, C, Morrell, LJ, Croft, DP, Dyer, JRG, Ioannou, C & Krause, J. (2007) A cost of leadership in human groups. Ethology 113: 821–824 [link]
  55. Morrell, LJ, Hunt, KL, Croft, DP & Krause, J. (2007) Diet, familiarity and shoaling decisions in guppies. Animal Behaviour 74: 311-319 [link]
  56. 2003-2006

  57. Kokko, H, Gunnarsson, TG, Morrell, LJ & Gill, JA. (2006) Why do female migratory birds arrive later than males? Journal of Animal Ecology 75: 1293-1303 [link]
  58. Croft, DP, Morrell, LJ, Wade, AS, Piyapong, C, Ioannou, CC, Dyer, JRG, Chapman, BB, Wong, Y & Krause, J. (2006) Predation risk as a driving force for sexual segregation: a cross-population comparison. American Naturalist 167: 867-887 [link]
  59. Kokko, H, López-Sepulcre, A & Morrell, LJ. (2006) From hawks and doves to self-consistent games of territorial behavior. American Naturalist 167: 901-912 [link]
  60. Morrell, LJ, Backwell, PRY & Metcalfe NB (2005) Fighting in fiddler crabs Uca mjoebergi: what determines duration? Animal Behaviour 70: 653-662 [link]
  61. Morrell, LJ, Lindstrom, J & Ruxton, GD (2005) Why are small males aggressive? Proceedings of the Royal Society of London Series B 272: 1235-1241 [link]
  62. Kokko, H & Morrell, LJ (2005) Mate guarding, male attractiveness and paternity under social monogamy. Behavioral Ecology 16: 724-731 [link]
  63. Morrell, LJ & Kokko, H (2005) Bridging the gap between mechanistic and adaptive explanations of territoriality. Behavioral Ecology and Sociobiology 57: 381-390 [link]
  64. Morrell, LJ (2004) Are behavioural trade-offs all they seem? Counter-intuitive resolution of the conflict between two behaviours. Behavioral Ecology and Sociobiology 56: 539-545 [link]
  65. Morrell, LJ & Kokko, H (2004) Can too strong female choice deteriorate male ornamentation? Proceedings of the Royal Society of London Series B 271: 1597-1604 [link]
  66. Morrell, LJ & Kokko, H (2003) Adaptive strategies of territory formation.  Behavioral Ecology and Sociobiology 54: 385-395 [link]

Other publications

  1. Morrell, LJ (2008) The sum of the parts. NERC Planet Earth Summer 2008: 31. [download]
  2. Compère, S, Morrell, LJ and Bousis, A. Once upon a time…Lygia. Environmental information leaflet. Igoumenitsa, Greece

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